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In terrestrial animals, plantigrade locomotion means walking with the toes and metatarsals flat on the ground. It is one of three forms of locomotion adopted by terrestrial mammals. The other options are digitigrade, walking on the toes with the heel and wrist permanently raised, and unguligrade, walking on the nail or nails of the toes with the heelwrist and the digits permanently raised. The leg of a plantigrade mammal includes the bones of the upper leg and lower leg . The leg of a digitigrade mammal also includes the metatarsalsmetacarpals, the bones that in a human compose the arch of the foot and the palm of the hand. The leg of an unguligrade mammal also includes the phalanges, the finger and toe bones.