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Tortoises are reptile species of the family Testudinidae of the order Testudines . They are particularly distinguished from turtles by being land-dwelling, while many turtle species are at least partly aquatic. However, like turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like other members of the suborder Cryptodira, they retract their necks and heads directly backwards into the shell to protect them.