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In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: The integument forming its outer layer, the nucellus , and female gametophyte in its center. The female gametophyte?specifically termed a megagametophyte?is also called the embryo sac in angiosperms. The megagametophyte produces an egg cell for the purpose of fertilization. After fertilization, the ovule develops into a seed.